Cuspicephalus

Cuspicephalus is an extinct genus of monofenestratan pterosaur known from Dorset in England.

Description
The skull is lightly built and has a length of 326 millimetres. It is very elongated, being just 55 millimetres high at the back, and has an triangular profile. Almost half of its length is accounted for by a large skull opening, the fenestra nasoantorbitalis, a confluence of the original fenestra antorbitalis with the bony nostrils. In front of this opening a low and elongated snout is present. The rostral index sensu Naish & Martill of this snout, in this case its length divided by its maximum height, is 5.4, the highest value known for any pterosaur. In the snout about eleven or twelve teeth are present; the total for the upper jaw is estimated at twenty-five to thirty. The teeth are largest in front and gradually decline in size towards the back. They have a broad base with an oval cross-section. Perhaps they are pointing outwards to a degree but the deformation of the fossil by compression makes this uncertain.

On top of the skull, behind the snout proper, above the fenestra a longitudinal low bony crest is present. It consists of fibrous bone, vertically directed, which might have formed the base for a much higher crest of soft tissue. The crest starts above the twelfth tooth position; its limit at the back is unknown because of damage but it is considered unlikely that it extended as far as the eyes, the roof of the skull not showing any trace of it. The crest is highest in the (preserved) middle.

Discovery
Cuspicephalus is known from the holotype MJML K1918, a partial skull which is missing the mandible and dentition, preserved on a single slab. It was collected in December 2009 by Steve Etches from Kimmeridge Bay on the Isle of Purbeck coast of Dorset. This locality is referred to the Autissiodorensis ammonite biozone of the lower part of the Kimmeridge Clay Formation, dating to the early Kimmeridgian stage of the Late Jurassic, about 155.7-153 million years ago.