Osteostraci

The class Osteostraci ("Bony Shields") was a group of bony-armored jawless fish, termed "ostracoderms", that lived in what is now North America, Europe and Russia from the Middle Silurian to Late Devonian.

Anatomically speaking, the osteostracans, especially the Devonian species, were among the most advanced of all known agnathans. This is due to the development of paired fins, and their complicated cranial anatomy. However, the osteostracans were more related to other agnathans than to jawed vertebrates as their inner ears are formed by two pairs of semicircular canals, similar to those of living lampreys, as opposed to the three pairs found in the inner ears of jawed vertebrates.

They were probably relatively good swimmers, possessing dorsal fins, paired pectoral fins, and a strong tail. The shield of bone covering the head formed a single piece, and so presumably did not grow during adult life. However, the way in which the bone was laid down makes it possible to examine the imprints of nerves and other soft tissues. This reveals the presence of complex sensory organs and the sides and upper surface of the head, which may have been used to sense vibrations, or could even have been electric organs.