Hyaenodon

Hyaenodon ("hyaena-toothed") is the type genus of Hyaenodontidae, a group of extinct carnivorous creodonts of the family Hyaenodontidae endemic to all continents except South America, Australia and Antarctica, living from 42—15.9 mya, existing for approximately 26.1 million years.[1] The various species of Hyaenodon competed with each other and with other hyaenodont genera (including Sinopa, Dissopsalis and Hyainailouros), and played important roles as predators in ecological communities as late as the Miocene in Africa and Asia.[2] Species of Hyaenodon have been shown to have successfully preyed on other large carnivores of their time, including a Dinictis nimravid "false saber-tooth cat" by analysis of tooth puncture marks on a fossil Dinictis skull in North Dakota.

Morphology
Some species of this genus were among the largest terrestrial carnivorous mammals of their time; others were only of the size of a marten. Hyaenodon is one of the longest lived genera of Hyaenodontidae, and is known from the Late Eocene to Early Miocene. Remains of many species are known from North America, Europe, Asia and Africa (In 1993 42 species were distinguished).[4]

Typical of early carnivorous mammals, individuals of Hyaenodon had a very massive skull but only a small brain. The skull is long with a narrow snout - much larger in relation to the length of the skull than in canine carnivores, for instance. The neck was shorter than the skull, while the body was long and robust and terminated in a long tail. Despite the name, these creatures are not related to hyenas.

The average weight of adult or subadult H. horridus, the largest North American species, is estimated to about 40 kilograms (88 lb) and may not have exceeded 60 kilograms (130 lb). H. gigas, the largest Hyaenodon species was much larger, being 500 kilograms (1,100 lb) and around 10 feet.[5][verification needed] H. crucians from the early Oligocene of North America is estimated to only 10 to 25 kilograms (22 to 55 lb). H. microdon and H. mustelinus from the late Eocene of North America were even smaller and weighed probably about 5 kilograms (11 lb).[6]

Compared to the generally larger (but closely related) Hyainailouros, Hyaenodon dentition was geared more towards shearing meat and less towards bone-crushing.

Range and species
In North America the last Hyaenodon disappeared along with species like H. brevirostris in the late Oligocene. In Europe they had already vanished earlier in the Oligocene. From the Miocene in Africa there are three species (H. andrewsi, H. matthewi and H. pilgrimi) known, but none of these reached the dimensions of Asian species like H. gigas and H. weilini.