Cycnorhamphus

Cycnorhamphus (meaning "swan beak") is a genus of ctenchasmatoid pterodactyloid pterosaur from the Late Jurassic of France and Germany.[1] It is probably synonymous with the genus Gallodactylus.

History and classification
In 1855 a fossil in a plate of shale from the Tithonian, found near Nusplingen in Württemberg, holotype GPIT "Orig. Quenstedt 1855, Taf. 1", was named Pterodactylus suevicus by Friedrich August Quenstedt.[2] In 1870 Harry Govier Seeley assigned it to a new genus: Cycnorhamphus. In 1907 however, Felix Plieninger rejected this split, an opinion then shared by most paleontologists. In 1974 Jacques Fabre, when concluding that a new species found and named by him, Gallodactylus canjuersensis, was the same genus as P. suevicus, did not revive Cycnorhamphus, but judged that the latter name was unavailable because of mistakes in the diagnosis by Seeley. P. suevicus thus became Gallodactylus suevicus. However, in 1996 Christopher Bennett pointed out that such mistakes do not invalidate a name and that therefore Cycnorhamphus has priority, making Gallodactylus canjuersensis C. canjuersensis.[2] In 2010 and 2012, Bennett published further re-studies of the fossils, concluding that the differences between the two species could be explained by age, sex or individual variation, and formally synonymized C. canjuersensis and C. suevicus.

Description
Cycnorhamphus had historically been assumed to have had long jaws with teeth at the very tip, akin to those of Pterodactylus antiquus. However, recent work on a specimen titled "The Painten Pelican"[4] has revealed that the animal possesses a very unusual jaw anatomy, with peg-like teeth at the jaw tips - blunter and stouter in older individuals -, jaw curvatures behind said teeth that form angled arcs away from the biting surface, forming thus an opening, and two poorly understood soft tissue structures occupying this opening from the upper jaw, showing mineralisation. The purpose of these adaptations is unknown,[5] but they are more obvious and well developed in adult animals. It has been speculated that a function akin to that of openbill storks, allowing the animal to hold hard invertebrates like mollusks and either crush or bisect them.