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The horse is a hoofed (ungulate) mammal, a subspecies of one of seven extant species of the family Equidae. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature into the large, single-toed animal of today.

Taxonomy and evolution[]

Mesohippus

Mesohippus, an ancestor of the modern horse.

Main article: Evolution of the horse

The horse as it is known today adapted by evolution to survive in areas of wide-open terrain with sparse vegetation, surviving in an ecosystem where other large grazing animals, especially ruminants, could not.[1] Horses and other equids are odd-toed ungulates of the order Perissodactyla, a group of mammals that was dominant during the Tertiary period. In the past, this order contained 14 families and many species, but only three families—Equidae (the horse and related species), the tapir and the rhinoceros—containing 18 known species have survived to the present day.[2] The earliest known member of the Equidae family was the Hyracotherium, which lived between 45 and 55 million years ago, during the Eocene period and had 4 toes on each front foot, and 3 toes on each back foot.[3] The extra toe on the front feet soon disappeared with the Mesohippus, which lived 32 to 37 million years ago,[4] and by about 5 million years ago, the modern Equus had developed.[5] The extra side toes shrank in size until they vanished. All that remains of them in modern horses is a set of small vestigial bones on the leg above the hoof,[6] known informally as ergots, chestnuts, or splint bones.[7] Their legs also lengthened as their toes disappeared and until they were a hoofed animal capable of running at great speed.[6]

Over millions of years, equid teeth also evolved from browsing on soft, tropical plants to adapt to browsing of drier plant material, and grazing of tougher plains grasses. Thus the proto-horses changed from leaf-eating forest-dwellers to grass-eating inhabitants of semi-arid regions worldwide, including the steppes of Eurasia and the Great Plains of North America.

About 15,000 years ago Equus ferus was a widespread, holarctic species. Horse bones from this time period, the late Pleistocene, are found in Europe, Eurasia, Beringia, and North America.[8] But by 10,000 years ago, the horse became extinct in North America and rare elsewhere.[9][10] The reasons for this extinction are not fully known, but one theory notes that extinction was contemporary with human arrival.[11] Another theory points to climate change, noting that approximately 12,500 years ago, the grasses characteristic of a steppe ecosystem gave way to shrub tundra, which was covered with unpalatable plants.[12]

References[]

  1. ^ Budiansky The Nature of Horses p. 31
  2. ^ Myers, Phil. "Order Perissodactyla". Animal Diversity Web. University of Michigan. http://animaldiversity.ummz.umich.edu/site/accounts/information/Perissodactyla.html. Retrieved on 2008-07-09. 
  3. ^ "Hyracotherium". Fossil Horses in Cyberspace. Florida Museum of Natural History. http://www.flmnh.ufl.edu/fhc/hyraco1.htm. Retrieved on 2008-07-09. 
  4. ^ "Mesohippus". Fossil Horses in Cyberspace. Florida Museum of Natural History. http://www.flmnh.ufl.edu/fhc/mesoh1.htm. Retrieved on 2008-07-09. 
  5. ^ "Equus". Florida Museum of Natural History. http://www.flmnh.ufl.edu/fhc/equus1.htm. Retrieved on 2008-07-09work= Fossil Horses in Cyberspace. 
  6. ^ a b "The Evolution of Horses". The Horse. American Museum of Natural History. http://www.amnh.org/exhibitions/horse/?section=evolution&page=evolution_b. Retrieved on 2008-07-09. 
  7. ^ Miller, Understanding the Ancient Secrets of the Horse's Mind, p. 20
  8. ^ Weinstock, J.; et al. (2005). "Evolution, systematics, and phylogeography of Pleistocene horses in the New World: a molecular perspective". PLoS Biology 3 (8): e241. doi:10.1371/journal.pbio.0030241. http://biology.plosjournals.org/perlserv/?request=get-document&doi=10.1371%2Fjournal.pbio.0030241&ct=1. Retrieved on 2008-12-19. 
  9. ^ Vila, C.; et al. (2001). "Widespread Origins of Domestic Horse Lineages" (PDF). Science (journal) 291: 474. doi:10.1126/science.291.5503.474. PMID 11161199. http://www.uky.edu/Ag/Horsemap/Maps/VILA.PDF. 
  10. ^ Luís, Cristina; et al. (2006). "Iberian Origins of New World Horse Breeds". Quaternary Science Reviews 97 (2): 107–113. doi:10.1093/jhered/esj020. PMID 16489143. http://jhered.oxfordjournals.org/cgi/content/full/97/2/107. 
  11. ^ Buck, Caitlin E.; Bard, Edouard (2007). "A calendar chronology for Pleistocene mammoth and horse extinction in North America based on Bayesian radiocarbon calibration". Quaternary Science Reviews 26 (17-18): 2031. doi:10.1016/j.quascirev.2007.06.013. 
  12. ^ LeQuire, Elise (2004-01-04). ""No Grass, No Horse"". http://www.thehorse.com/ViewArticle.aspx?ID=4849. 
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